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Fraxinus (), commonly called ash, is a of plants in the olive and lilac family, ,

(1995). 9780376038500, .
and comprises 45–65 species of usually medium-to-large , most of which are (dropping their leaves in autumn), although some species are . The genus is widespread throughout much of Europe, Asia, and North America.

The leaves are usually , and mostly (divided into leaflets in a feather-like arrangement). The seeds, known as "keys", are botanically fruits of the type called samara. Some species are , having male and female flowers on separate plants.

Ash wood is strong and elastic, and used for the handles of tools. Musical instrument makers use it for and for . The Morgan Motor Company makes the frames of sports cars from ash wood. In , the were the of ash trees.


Etymology
The tree's common English name, "ash", derives from the , from the Proto-Indo-European name for the tree, while the name of the genus originated in , from a Proto-Indo-European word for . Both words also meant "", as ash wood was used for shafts.
(1997). 9781884964985, Taylor & Francis. .


Description
The leaves of ash trees are usually (rarely in whorls), and mostly . The seeds, known as "keys", are botanically single-winged fruits of the type called samara. Most Fraxinus species are , having male and female flowers on separate plants. The male flowers have two stamens. If a calyx is present, it has four lobes; if there is a corolla, it has four lobes or four petals, which are white or pale yellow.

File:NarrowleafAsh.jpg|Leafy shoot of F. angustifolia File:Ash flower.JPG|Flowers of F. excelsior File:EurAshSeeds.jpg|Winged fruits (samaras) of F. excelsior


Evolution

Fossil history
The oldest fossils that are clearly Fraxinus are from the (49–39 million years ago) of southeast North America, including the extinct species F. wilcoxiana. Fossil pollen of F. angustifolia is known from the (12 million years ago) of Europe. F. oishii winged fruits have been found in the of Korea.


Taxonomy
The genus Fraxinus was described by in 1753. The name remains accepted by taxonomists. Multiple authors have described other tree genera that are synonymous with Fraxinus: Ornus by the German botanist and physician Georg Rudolf Boehmer in 1760; Fraxinoides by the German physician Friedrich Kasimir Medikus in 1791; Mannaphorus by the French polymath Constantine Samuel Rafinesque in 1818; Calycomelia by the Czech in 1834; Leptalix, Ornanthes, Samarpsea (misspelt) and Samarpses, Apilia and Aplilia by Rafinesque, all in 1838; Meliopsis by the German botanist Ludwig Reichenbach in 1841; and Petlomelia by the Belgian priest in 1914.


External phylogeny
The Angiosperm Phylogeny Group has classified Fraxinus as part of the (the family of flowering woody plants), within the order (the order, including many aromatic herbs).


Internal phylogeny
Species are arranged into sections identified by phylogenetic analysis of clades within the Fraxinus genus:

Section Dipetalae
  • Torr. ex S.Watson – singleleaf ash
  • Fraxinus dipetala Hook. & Arn. – California ash or two-petal ash
  • Moran – Chaparral ash
  • Fraxinus quadrangulata Michx. – blue ash
  • Fraxinus trifoliolata

Section Fraxinus
  • Fraxinus angustifolia Vahl – narrow-leaved ash
    • Fraxinus angustifolia subsp. oxycarpa – Caucasian ash
    • Fraxinus angustifolia subsp. syriaca
  • Fraxinus excelsior L. – European ash
  • Fraxinus mandschurica Rupr. – Manchurian ash
  • Marshall – black ash
  • Fraxinus pallisiae Wilmott – Pallis' ash
  • Fraxinus sogdiana – Tianshan ash

Section Melioides sensu lato
  • Fraxinus chiisanensis – Jirisan ash
  • Fraxinus cuspidata Torr. – fragrant ash
  • Fraxinus platypoda – Chinese red ash
  • Fraxinus spaethiana Lingelsh. – Späth's ash

Section Melioides sensu stricto
  • Fraxinus albicans Buckley – Texas ash
  • Fraxinus americana L. – white ash or American ash
  • Fraxinus berlandieriana DC. – Mexican ash
  • Fraxinus caroliniana Mill. – Carolina ash
  • Fraxinus latifolia Benth. – Oregon ash
  • Fraxinus papillosa Lingelsh. – Chihuahua ash
  • Fraxinus pennsylvanica Marshall – green ash
  • Fraxinus profunda (Bush) Bush – pumpkin ash
  • (Wenz.) Lingelsh. – Shamel ash or Tropical ash
  • Fraxinus velutina Torr. – velvet ash or Arizona ash

Section Ornus
  • Fraxinus apertisquamifera
  • Fraxinus baroniana
  • Fraxinus bungeana DC. – Bunge's ash
  • Fraxinus chinensis Roxb. – Chinese ash or Korean ash
  • Fraxinus floribunda Wall. – Himalayan manna ash
  • Fraxinus griffithii C.B.Clarke – Griffith's ash
  • Fraxinus insularis Hemsl. – Chinese flowering ash
  • Fraxinus japonica – Japanese ash
  • Fraxinus lanuginosa – Japanese ash
  • Fraxinus longicuspis
  • Fraxinus malacophylla
  • Fraxinus micrantha Lingelsh.
  • L. – manna ash or flowering ash
  • Lingelsh.
  • Fraxinus sieboldiana Blume – Japanese flowering ash

Section Pauciflorae

Section Sciadanthus
  • Fraxinus dimorpha
  • Fraxinus hubeiensis Ch'u & Shang & Su – 湖北梣, Hubei qin
  • Fraxinus xanthoxyloides (G.Don) Wall. ex DC. – Afghan ash


Ecology and distribution
The genus Fraxinus is widespread throughout much of Europe, Asia, and North America. The genus is primarily temperate or subtropical; 22 of the species occur in China, while for example Italy has 4 species. Both native and introduced Fraxinus species occur in almost every contiguous state of the United States and all the southern provinces of Canada.

Ash species provide habitat and food for the larvae of many insects including , plant bugs, , , and , as well as birds and mammals.

The emerald ash borer ( Agrilus planipennis), is a wood-boring accidentally introduced to North America from eastern Asia via solid wood packing material in the late 1980s to early 1990s. It has killed tens of millions of trees in 22 states in the United States and neighbouring and in Canada. It threatens some seven billion ash trees in North America. Three native Asian wasp species, natural predators of the beetle, have been evaluated as possible biological controls. The public was cautioned to avoid transporting unfinished wood products, such as firewood, to slow the spread of the pest. Damage occurs when emerald ash borer larvae feed on the inner bark, of ash trees, preventing nutrient and water transportation.

The European ash, Fraxinus excelsior, has been affected by the Hymenoscyphus fraxineus, causing chalara in a large number of trees since the mid-1990s, particularly in eastern and northern Europe. The disease has infected about 90% of Denmark's ash trees. In 2012 in the UK, ash dieback was found in mature woodland. BBC News 'Ash dieback' fungus, Chalara fraxinea found in UK countryside. Retrieved 25 October 2012. The combination of emerald ash borer and ash dieback has threatened ash populations in Europe, but trees in mixed landscapes appear to have some resistance to the disease.

File:Agrilus planipennis 001.jpg|Emerald ash borer
adult File:Eablarva.jpg|Emerald ash borer File:Bore remnants.jpg|Emerald ash borer damage on a fallen trunk File:Chalara ash dieback - symptoms - 39.jpg|Chalara ash dieback File:Canker on Ash.JPG| on an ash tree


Uses
Ash is a and is dense, around 670 kg/m3 for Fraxinus americana, the white ash, and 710 kg/m3 for Fraxinus excelsior, the European ash. The wood of the European ash is strong and elastic, making it suitable for uses such as the handles of tools. It is a good .

The Fender musical instrument company has used ash as a for its since 1950. Species used for guitar building include . SWAMP ASH Lumber Guide: 8/4 Lightweight Guitar Wood 2020 15 December 2018 www.commercialforestproducts.com, accessed 27 September 2020 Ash is in addition used for making . It has been described as resonant, providing a balanced tone for both high and low notes. Ash wood can be used for furniture, agricultural tools, and household objects such as bowls, candlesticks, and spoons. The Morgan Motor Company of Great Britain still manufactures sports cars with frames made from ash.

The green ash ( F. pennsylvanica) is widely planted as a street tree in the United States. The inner bark of the blue ash ( F. quadrangulata) has been used as a source for blue .Oklahoma Biological Survey: Fraxinus quadrangulata In Sicily, Italy, a sugary is obtained from the resinous sap of the , extracted by making cuts in the bark. The young seedpods of the European ash, known as "keys", are edible; in Britain, they are traditionally pickled with vinegar, sugar and spices. A range of pharmacologically active compounds exist in Fraxinus species, with for example anti-inflammatory, , and antihyperglycaemic properties which might find practical applications.

File:Fender Telecaster lite ash headstock (2005-11-27 07.55.40 by Pierre Journel).jpg|Ash headstock of a Fender Telecaster
electric guitar File:Shovel 24cm (cropped).jpg|Ash is widely used for tool handles. File:Hurling Ball and Hurley.JPG|Hurley (playing stick used in and )

File:Flamed Quartersawn Ash Guitar Top.jpg|5/16" thick flame figure ash top, unmilled File:Ash_Table_by_Ben_Barclay.jpg|Ash coffee table File:MorganWoodDoorFrame.jpg|Ash frame of a
Morgan Motor Company car


Mythology and folklore
In , the , their name meaning "ash trees", are associated with the ash, perhaps specifically of the manna ash ( ), as were nymphs associated with the . They appear in 's , which states that they were born when drops of Ouranos's blood fell on the earth (Gaia).Caldwell, Richard, Hesiod's Theogony, Focus Publishing/R. Pullins Company (June 1, 1987). . p. 38 n. 178–187: "The nymphs called Meliai are properly "ash-tree" nymphs; the Greek word for ash-trees is meliai also". In , a vast, evergreen ash tree ("the steed () of "), watered by three magical springs, serves as , sustaining the nine worlds of the in its roots and branches. Askr, the first man in Norse myth, literally means 'ash'.
(2026). 9780859915137, D.S. Brewer.
In , an ash stake could be used to kill a .
(2026). 9781493138388, Xlibris Corporation.
In the Old English Latin alphabet, Æ was the letter æsc, "ash tree". This transliterated the Anglo-Saxon futhorc rune ᚫ.

The Welsh Llwyn Onn, "The Ash Grove", sings in the English version by "The ash grove, how graceful, how plainly 'tis speaking; The lark through its branches is gazing on me".


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