Fraxinus (), commonly called ash, is a genus of plants in the olive and lilac family, Oleaceae, and comprises 45–65 species of usually medium-to-large , most of which are deciduous (dropping their leaves in autumn), although some Subtropics species are evergreen. The genus is widespread throughout much of Europe, Asia, and North America.
The leaves are usually opposite leaves, and mostly pinnate (divided into leaflets in a feather-like arrangement). The seeds, known as "keys", are botanically fruits of the type called samara. Some species are Dioecy, having male and female flowers on separate plants.
Ash wood is strong and elastic, and used for the handles of tools. Musical instrument makers use it for and for . The Morgan Motor Company makes the frames of sports cars from ash wood. In Greek mythology, the Meliae were the of ash trees.
Etymology
The tree's common English name, "ash", derives from the
Old English , from the Proto-Indo-European name for the tree, while the name of the genus originated in Latin , from a Proto-Indo-European word for
birch. Both words also meant "
spear", as ash wood was used for shafts.
Description
The leaves of ash trees are usually
opposite leaves (rarely in whorls), and mostly
pinnate. The seeds, known as "keys", are botanically single-winged fruits of the type called samara. Most
Fraxinus species are
dioecy, having male and female flowers on separate plants. The male flowers have two stamens. If a calyx is present, it has four lobes; if there is a corolla, it has four lobes or four petals, which are white or pale yellow.
File:NarrowleafAsh.jpg|Leafy shoot of F. angustifolia
File:Ash flower.JPG|Flowers of F. excelsior
File:EurAshSeeds.jpg|Winged fruits (samaras) of F. excelsior
Evolution
Fossil history
The oldest fossils that are clearly
Fraxinus are from the
Middle Eocene (49–39 million years ago) of southeast North America, including the extinct species
F. wilcoxiana.
Fossil pollen of
F. angustifolia is known from the
Upper Miocene (12 million years ago) of Europe.
F. oishii winged fruits have been found in the
Middle Miocene of Korea.
Taxonomy
The genus
Fraxinus was described by
Carl Linnaeus in 1753. The name remains accepted by taxonomists.
Multiple authors have described other tree genera that are synonymous with
Fraxinus:
Ornus by the German botanist and physician Georg Rudolf Boehmer in 1760;
Fraxinoides by the German physician Friedrich Kasimir Medikus in 1791;
Mannaphorus by the French polymath Constantine Samuel Rafinesque in 1818;
Calycomelia by the Czech in 1834;
Leptalix,
Ornanthes,
Samarpsea (misspelt) and
Samarpses,
Apilia and
Aplilia by Rafinesque, all in 1838;
Meliopsis by the German botanist Ludwig Reichenbach in 1841; and
Petlomelia by the Belgian priest
Julius Nieuwland in 1914.
External phylogeny
The Angiosperm Phylogeny Group has classified
Fraxinus as part of the
Oleaceae (the
olive family of flowering woody plants), within the order
Lamiales (the
Mentha order, including many aromatic herbs).
Internal phylogeny
Species are arranged into sections identified by phylogenetic analysis of clades within the
Fraxinus genus:
- Section Dipetalae
-
Fraxinus anomala Torr. ex S.Watson – singleleaf ash
-
Fraxinus dipetala Hook. & Arn. – California ash or two-petal ash
-
Fraxinus parryi Moran – Chaparral ash
-
Fraxinus quadrangulata Michx. – blue ash
-
Fraxinus trifoliolata
- Section Fraxinus
-
Fraxinus angustifolia Vahl – narrow-leaved ash
-
Fraxinus angustifolia subsp. oxycarpa – Caucasian ash
-
Fraxinus angustifolia subsp. syriaca
-
Fraxinus excelsior L. – European ash
-
Fraxinus mandschurica Rupr. – Manchurian ash
-
Fraxinus nigra Marshall – black ash
-
Fraxinus pallisiae Wilmott – Pallis' ash
-
Fraxinus sogdiana – Tianshan ash
- Section Melioides sensu lato
-
Fraxinus chiisanensis – Jirisan ash
-
Fraxinus cuspidata Torr. – fragrant ash
-
Fraxinus platypoda – Chinese red ash
-
Fraxinus spaethiana Lingelsh. – Späth's ash
- Section Melioides sensu stricto
-
Fraxinus albicans Buckley – Texas ash
-
Fraxinus americana L. – white ash or American ash
-
Fraxinus berlandieriana DC. – Mexican ash
-
Fraxinus caroliniana Mill. – Carolina ash
-
Fraxinus latifolia Benth. – Oregon ash
-
Fraxinus papillosa Lingelsh. – Chihuahua ash
-
Fraxinus pennsylvanica Marshall – green ash
-
Fraxinus profunda (Bush) Bush – pumpkin ash
-
Fraxinus uhdei (Wenz.) Lingelsh. – Shamel ash or Tropical ash
-
Fraxinus velutina Torr. – velvet ash or Arizona ash
- Section Ornus
-
Fraxinus apertisquamifera
-
Fraxinus baroniana
-
Fraxinus bungeana DC. – Bunge's ash
-
Fraxinus chinensis Roxb. – Chinese ash or Korean ash
-
Fraxinus floribunda Wall. – Himalayan manna ash
-
Fraxinus griffithii C.B.Clarke – Griffith's ash
-
Fraxinus insularis Hemsl. – Chinese flowering ash
-
Fraxinus japonica – Japanese ash
-
Fraxinus lanuginosa – Japanese ash
-
Fraxinus longicuspis
-
Fraxinus malacophylla
-
Fraxinus micrantha Lingelsh.
-
Fraxinus ornus L. – manna ash or flowering ash
-
Fraxinus paxiana Lingelsh.
-
Fraxinus sieboldiana Blume – Japanese flowering ash
- Section Pauciflorae
- Section Sciadanthus
-
Fraxinus dimorpha
-
Fraxinus hubeiensis Ch'u & Shang & Su – 湖北梣, Hubei qin
-
Fraxinus xanthoxyloides (G.Don) Wall. ex DC. – Afghan ash
Ecology and distribution
The genus
Fraxinus is widespread throughout much of Europe, Asia, and North America.
The genus is primarily temperate or subtropical; 22 of the species occur in China,
while for example Italy has 4 species.
Both native and introduced
Fraxinus species occur in almost every contiguous state of the United States and all the southern provinces of Canada.
Ash species provide habitat and food for the larvae of many insects including , plant bugs, , , and , as well as birds and mammals.
The emerald ash borer ( Agrilus planipennis), is a wood-boring beetle accidentally introduced to North America from eastern Asia via solid wood packing material in the late 1980s to early 1990s. It has killed tens of millions of trees in 22 states in the United States and neighbouring Ontario and Quebec in Canada. It threatens some seven billion ash trees in North America. Three native Asian wasp species, natural predators of the beetle, have been evaluated as possible biological controls. The public was cautioned to avoid transporting unfinished wood products, such as firewood, to slow the spread of the pest. Damage occurs when emerald ash borer larvae feed on the inner bark, phloem of ash trees, preventing nutrient and water transportation.
The European ash, Fraxinus excelsior, has been affected by the fungus Hymenoscyphus fraxineus, causing chalara ash dieback in a large number of trees since the mid-1990s, particularly in eastern and northern Europe. The disease has infected about 90% of Denmark's ash trees. In 2012 in the UK, ash dieback was found in mature woodland.[ BBC News 'Ash dieback' fungus, Chalara fraxinea found in UK countryside. Retrieved 25 October 2012.] The combination of emerald ash borer and ash dieback has threatened ash populations in Europe, but trees in mixed landscapes appear to have some resistance to the disease.
File:Agrilus planipennis 001.jpg|Emerald ash borer
adult
File:Eablarva.jpg|Emerald ash borer larva
File:Bore remnants.jpg|Emerald ash borer damage on a fallen trunk
File:Chalara ash dieback - symptoms - 39.jpg|Chalara ash dieback
File:Canker on Ash.JPG|Canker on an ash tree
Uses
Ash is a
hardwood and is dense, around 670 kg/m
3 for
Fraxinus americana, the white ash,
and 710 kg/m
3 for
Fraxinus excelsior, the European ash.
The wood of the European ash is strong and elastic, making it suitable for uses such as the handles of tools.
It is a good
firewood.
The Fender musical instrument company has used ash as a tonewood for its since 1950. Species used for guitar building include swamp ash.[ SWAMP ASH Lumber Guide: 8/4 Lightweight Guitar Wood 2020 15 December 2018 www.commercialforestproducts.com, accessed 27 September 2020] Ash is in addition used for making . It has been described as resonant, providing a balanced tone for both high and low notes. Ash wood can be used for furniture, agricultural tools, and household objects such as bowls, candlesticks, and spoons. The Morgan Motor Company of Great Britain still manufactures sports cars with frames made from ash.
The green ash ( F. pennsylvanica) is widely planted as a street tree in the United States. The inner bark of the blue ash ( F. quadrangulata) has been used as a source for blue dye.[Oklahoma Biological Survey: Fraxinus quadrangulata ] In Sicily, Italy, a sugary manna is obtained from the resinous sap of the manna ash, extracted by making cuts in the bark. The young seedpods of the European ash, known as "keys", are edible; in Britain, they are traditionally pickled with vinegar, sugar and spices. A range of pharmacologically active compounds exist in Fraxinus species, with for example anti-inflammatory, antihypertensive, and antihyperglycaemic properties which might find practical applications.
File:Fender Telecaster lite ash headstock (2005-11-27 07.55.40 by Pierre Journel).jpg|Ash headstock of a Fender Telecaster
electric guitar
File:Shovel 24cm (cropped).jpg|Ash is widely used for tool handles.
File:Hurling Ball and Hurley.JPG|Hurley (playing stick used in hurling and camogie)
File:Flamed Quartersawn Ash Guitar Top.jpg|5/16" thick flame figure quartersawn ash guitar top, unmilled
File:Ash_Table_by_Ben_Barclay.jpg|Ash coffee table
File:MorganWoodDoorFrame.jpg|Ash frame of a
Morgan Motor Company car
Mythology and folklore
In
Greek mythology, the
Meliae, their name meaning "ash trees", are
associated with the ash, perhaps specifically of the manna ash (
Fraxinus ornus), as
were nymphs associated with the
oak. They appear in
Hesiod's
Theogony, which states that they were born when drops of Ouranos's blood fell on the earth (Gaia).
[Caldwell, Richard, Hesiod's Theogony, Focus Publishing/R. Pullins Company (June 1, 1987). . p. 38 n. 178–187: "The nymphs called Meliai are properly "ash-tree" nymphs; the Greek word for ash-trees is meliai also".]
In
Norse mythology, a vast, evergreen ash tree
Yggdrasil ("the steed (
gallows) of
Odin"), watered by three magical springs, serves as
axis mundi, sustaining the nine worlds of the
cosmos in its roots and branches.
Askr, the first man in Norse myth, literally means 'ash'.
In
Slavic folklore, an ash stake could be used to kill a
vampire.
In the Old English Latin alphabet, Æ was the letter æsc, "ash tree".
This transliterated the Anglo-Saxon futhorc rune ᚫ.
The Welsh folk song Llwyn Onn, "The Ash Grove", sings in the English version by John Oxenford "The ash grove, how graceful, how plainly 'tis speaking; The lark through its branches is gazing on me".
External links